Can the relationship between beef color and tenderness be explained by myoglobin regulating calpain-1?

Can the relationship between beef color and tenderness be explained by myoglobin regulating calpain-1?
Jade V. Cooper, Surendranath P. Suman, Zachary D. Callahan, Karl C. Kerns, Michal Zigo, Peter Sutovsky, Steven M. Lonergan, Carol L. Lorenzen